
The game also includes the GHTunes from World Tour where new songs can be created and shared. Performances are rated with stars and a certain amount is needed to progress and unlock a new set.
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Progress is made by completing sets of songs, rather than a series of gigs as in Guitar Hero: World Tour, set in venues that were in important in the band's career. Bonus features include various videos and trivia about the songs.

The music selection consists of 28 master recordings and 21 additional songs by others selected by the band. The members also did motion capture sessions for the avatars in the game. This edition adds an Expert+ difficulty mode where players need to drum using a second bass pedal to mimic the playing style of Lars Ulrich, the drummer of the band. The storyline describes how a band wants to follow in the footsteps of the Metallica and become a leading act on their tour. This title largely borrows from the novelties introduced in Guitar Hero: World Tour, with band career modes for single or multiple players with a storyline drawn as a cartoon, competitive modes online, and the possibility to play with different instruments such as a regular guitar, a bass guitar, drums, and doing the vocals. Unleash the true power of infection raging within you and kill Alex Mercer. The virus has left the city in ruins, but it's not going to stop you from achieving your goal. Choose from a wide variety of new Mutation perks, including better locomotion abilities, sharper claws and enhanced disguise abilities, to build and customize the ultimate Prototype. Track down your enemies by emitting sonar pulses from your body. Wield the unbelievable Tendril power that allows you to lift and tear apart cars and military vehicles, ensnare powerful infected beasts and shred groups of enemies to pieces. James Heller, and hunt, kill and become anybody who stands in your way to absorb memories and gain new abilities. Take on swarms of highly trained Blackwatch soldiers. Witness building-size infected beasts ambush entire city streets.

Cut through the wastelands of post-viral New York Zero and build a genetic arsenal of all-new biological weapons and abilities on your quest to murder your maker and the devil himself, Alex Mercer.

Become the ultimate shape-shifting weapon in Prototype 2.
